diff --git a/src/OpenGL/compiler/OutputASM.cpp b/src/OpenGL/compiler/OutputASM.cpp
index 9ed34b2..c18ecf7 100644
--- a/src/OpenGL/compiler/OutputASM.cpp
+++ b/src/OpenGL/compiler/OutputASM.cpp
@@ -400,12 +400,12 @@
 		registerIndex = 0;
 	}
 
-	Attribute::Attribute(GLenum type, const std::string &name, int arraySize, int location, int registerIndex)
+	Attribute::Attribute(GLenum type, const std::string &name, int arraySize, int layoutLocation, int registerIndex)
 	{
 		this->type = type;
 		this->name = name;
 		this->arraySize = arraySize;
-		this->location = location;
+		this->layoutLocation = layoutLocation;
 		this->registerIndex = registerIndex;
 	}
 
diff --git a/src/OpenGL/compiler/OutputASM.h b/src/OpenGL/compiler/OutputASM.h
index 2f90266..b037a67 100644
--- a/src/OpenGL/compiler/OutputASM.h
+++ b/src/OpenGL/compiler/OutputASM.h
@@ -144,12 +144,12 @@
 	struct Attribute
 	{
 		Attribute();
-		Attribute(GLenum type, const std::string &name, int arraySize, int location, int registerIndex);
+		Attribute(GLenum type, const std::string &name, int arraySize, int layoutLocation, int registerIndex);
 
 		GLenum type;
 		std::string name;
 		int arraySize;
-		int location;
+		int layoutLocation;
 
 		int registerIndex;
 	};
diff --git a/src/OpenGL/libGLESv2/Program.cpp b/src/OpenGL/libGLESv2/Program.cpp
index 77912cc..dc64813 100644
--- a/src/OpenGL/libGLESv2/Program.cpp
+++ b/src/OpenGL/libGLESv2/Program.cpp
@@ -277,19 +277,7 @@
 
 	GLint Program::getAttributeLocation(const char *name)
 	{
-		if(name)
-		{
-			std::string strName(name);
-			for(auto const &it : linkedAttribute)
-			{
-				if(it.name == strName)
-				{
-					return getAttributeBinding(it);
-				}
-			}
-		}
-
-		return -1;
+		return name ? getAttributeLocation(std::string(name)) : -1;
 	}
 
 	int Program::getAttributeStream(int attributeIndex)
@@ -1591,80 +1579,55 @@
 	// Determines the mapping between GL attributes and vertex stream usage indices
 	bool Program::linkAttributes()
 	{
+		static_assert(MAX_VERTEX_ATTRIBS <= 32, "attribute count exceeds bitfield count");
 		unsigned int usedLocations = 0;
 
-		// Link attributes that have a binding location
+		// Link attributes that have a GLSL layout location qualifier
 		for(auto const &attribute : vertexShader->activeAttributes)
 		{
-			int location = (attributeBinding.find(attribute.name) != attributeBinding.end()) ? attributeBinding[attribute.name] : -1;
-
-			if(location != -1)   // Set by glBindAttribLocation
+			if(attribute.layoutLocation != -1)
 			{
-				int rows = VariableRegisterCount(attribute.type);
-
-				if(rows + location > MAX_VERTEX_ATTRIBS)
+				if(!linkAttribute(attribute, attribute.layoutLocation, usedLocations))
 				{
-					appendToInfoLog("Active attribute (%s) at location %d is too big to fit", attribute.name.c_str(), location);
 					return false;
 				}
-
-				// In GLSL 3.00, attribute aliasing produces a link error
-				// In GLSL 1.00, attribute aliasing is allowed
-				if(vertexShader->getShaderVersion() >= 300)
-				{
-					for(auto const &it : linkedAttribute)
-					{
-						int itLocStart = getAttributeBinding(it);
-						ASSERT(itLocStart >= 0);
-						int itLocEnd = itLocStart + VariableRegisterCount(it.type);
-						for(int i = 0; i < rows; i++)
-						{
-							int loc = location + i;
-							if((loc >= itLocStart) && (loc < itLocEnd))
-							{
-								appendToInfoLog("Attribute '%s' aliases attribute '%s' at location %d", attribute.name.c_str(), it.name.c_str(), location);
-								return false;
-							}
-						}
-					}
-				}
-
-				linkedAttributeLocation[attribute.name] = location;
-				linkedAttribute.push_back(attribute);
-				for(int i = 0; i < rows; i++)
-				{
-					usedLocations |= 1 << (location + i);
-				}
 			}
 		}
 
-		// Link attributes that don't have a binding location
+		// Link attributes that have an API provided binding location but no GLSL layout location
 		for(auto const &attribute : vertexShader->activeAttributes)
 		{
-			int location = (attributeBinding.find(attribute.name) != attributeBinding.end()) ? attributeBinding[attribute.name] : -1;
+			int bindingLocation = (attributeBinding.find(attribute.name) != attributeBinding.end()) ? attributeBinding[attribute.name] : -1;
 
-			if(location == -1)   // Not set by glBindAttribLocation
+			if(attribute.layoutLocation == -1 && bindingLocation != -1)
 			{
-				int rows = VariableRegisterCount(attribute.type);
-				int availableIndex = AllocateFirstFreeBits(&usedLocations, rows, MAX_VERTEX_ATTRIBS);
-
-				if(availableIndex == -1 || availableIndex + rows > MAX_VERTEX_ATTRIBS)
+				if(!linkAttribute(attribute, bindingLocation, usedLocations))
 				{
-					appendToInfoLog("Too many active attributes (%s)", attribute.name.c_str());
-					return false;   // Fail to link
+					return false;
 				}
-
-				linkedAttributeLocation[attribute.name] = availableIndex;
-				linkedAttribute.push_back(attribute);
 			}
 		}
 
-		for(auto const &it : linkedAttribute)
+		// Link attributes that don't have a binding location nor a layout location
+		for(auto const &attribute : vertexShader->activeAttributes)
 		{
-			int location = getAttributeBinding(it);
+			if(attribute.layoutLocation == -1 && attributeBinding.find(attribute.name) == attributeBinding.end())
+			{
+				if(!linkAttribute(attribute, -1, usedLocations))
+				{
+					return false;
+				}
+			}
+		}
+
+		ASSERT(linkedAttribute.size() == vertexShader->activeAttributes.size());
+
+		for(auto const &attribute : linkedAttribute)
+		{
+			int location = getAttributeLocation(attribute.name);
 			ASSERT(location >= 0);
-			int index = vertexShader->getSemanticIndex(it.name);
-			int rows = std::max(VariableRegisterCount(it.type), 1);
+			int index = vertexShader->getSemanticIndex(attribute.name);
+			int rows = VariableRegisterCount(attribute.type);
 
 			for(int r = 0; r < rows; r++)
 			{
@@ -1675,17 +1638,69 @@
 		return true;
 	}
 
-	int Program::getAttributeBinding(const glsl::Attribute &attribute)
+	bool Program::linkAttribute(const glsl::Attribute &attribute, int location, unsigned int &usedLocations)
 	{
-		if(attribute.location != -1)
+		int rows = VariableRegisterCount(attribute.type);
+
+		if(location == -1)
 		{
-			return attribute.location;
+			location = AllocateFirstFreeBits(&usedLocations, rows, MAX_VERTEX_ATTRIBS);
+
+			if(location == -1 || location + rows > MAX_VERTEX_ATTRIBS)
+			{
+				appendToInfoLog("Too many active attributes (%s)", attribute.name.c_str());
+				return false;   // Fail to link
+			}
+		}
+		else
+		{
+			if(rows + location > MAX_VERTEX_ATTRIBS)
+			{
+				appendToInfoLog("Active attribute (%s) at location %d is too big to fit", attribute.name.c_str(), location);
+				return false;
+			}
+
+			// In GLSL 3.00, attribute aliasing produces a link error
+			// In GLSL 1.00, attribute aliasing is allowed
+			if(vertexShader->getShaderVersion() >= 300)
+			{
+				for(auto const &previousAttrib : linkedAttribute)
+				{
+					int previousLocation = getAttributeLocation(previousAttrib.name);
+					int previousRows = VariableRegisterCount(previousAttrib.type);
+
+					if(location >= previousLocation && location < previousLocation + previousRows)
+					{
+						appendToInfoLog("Attribute '%s' aliases attribute '%s' at location %d", attribute.name.c_str(), previousAttrib.name.c_str(), location);
+						return false;
+					}
+
+					if(location <= previousLocation && location + rows > previousLocation)
+					{
+						appendToInfoLog("Attribute '%s' aliases attribute '%s' at location %d", attribute.name.c_str(), previousAttrib.name.c_str(), previousLocation);
+						return false;
+					}
+				}
+			}
+
+			for(int i = 0; i < rows; i++)
+			{
+				usedLocations |= 1 << (location + i);
+			}
 		}
 
-		std::map<std::string, GLuint>::const_iterator it = linkedAttributeLocation.find(attribute.name);
-		if(it != linkedAttributeLocation.end())
+		linkedAttributeLocation[attribute.name] = location;
+		linkedAttribute.push_back(attribute);
+
+		return true;
+	}
+
+	int Program::getAttributeLocation(const std::string &name)
+	{
+		std::map<std::string, GLuint>::const_iterator attribute = linkedAttributeLocation.find(name);
+		if(attribute != linkedAttributeLocation.end())
 		{
-			return it->second;
+			return attribute->second;
 		}
 
 		return -1;
diff --git a/src/OpenGL/libGLESv2/Program.h b/src/OpenGL/libGLESv2/Program.h
index 501d081..a232926 100644
--- a/src/OpenGL/libGLESv2/Program.h
+++ b/src/OpenGL/libGLESv2/Program.h
@@ -228,7 +228,8 @@
 		bool linkTransformFeedback();
 
 		bool linkAttributes();
-		int getAttributeBinding(const glsl::Attribute &attribute);
+		bool linkAttribute(const glsl::Attribute &attribute, int location, unsigned int &usedLocations);
+		int getAttributeLocation(const std::string &name);
 
 		Uniform *getUniform(const std::string &name) const;
 		bool linkUniforms(const Shader *shader);
diff --git a/src/OpenGL/libGLESv2/libGLESv2.cpp b/src/OpenGL/libGLESv2/libGLESv2.cpp
index 789b0c8..b3f1d20 100644
--- a/src/OpenGL/libGLESv2/libGLESv2.cpp
+++ b/src/OpenGL/libGLESv2/libGLESv2.cpp
@@ -2345,7 +2345,6 @@
 
 	if(context)
 	{
-
 		es2::Program *programObject = context->getProgram(program);
 
 		if(!programObject)
diff --git a/tests/unittests/unittests.cpp b/tests/unittests/unittests.cpp
index 80bc887..91b34cd 100644
--- a/tests/unittests/unittests.cpp
+++ b/tests/unittests/unittests.cpp
@@ -266,6 +266,8 @@
 		glGetProgramiv(ph.program, GL_LINK_STATUS, &linkStatus);
 		EXPECT_NE(linkStatus, 0);
 
+		EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
 		return ph;
 	}
 
@@ -274,6 +276,8 @@
 		glDeleteShader(ph.fragmentShader);
 		glDeleteShader(ph.vertexShader);
 		glDeleteProgram(ph.program);
+
+		EXPECT_GLENUM_EQ(GL_NONE, glGetError());
 	}
 
 	void drawQuad(GLuint program, const char* textureName = nullptr)
@@ -411,8 +415,6 @@
 {
 	Initialize(3, false);
 
-	unsigned char green[4] = { 0, 255, 0, 255 };
-
 	const std::string vs =
 		"#version 300 es\n"
 		"in vec4 position;\n"
@@ -464,6 +466,7 @@
 
 	deleteProgram(ph);
 
+	unsigned char green[4] = { 0, 255, 0, 255 };
 	expectFramebufferColor(green);
 
 	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
@@ -476,8 +479,6 @@
 {
 	Initialize(3, false);
 
-	unsigned char green[4] = { 0, 255, 0, 255 };
-
 	const std::string vs =
 		"#version 300 es\n"
 		"in vec4 position;\n"
@@ -521,6 +522,160 @@
 
 	deleteProgram(ph);
 
+	unsigned char green[4] = { 0, 255, 0, 255 };
+	expectFramebufferColor(green);
+
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	Uninitialize();
+}
+
+// Test vertex attribute location linking
+TEST_F(SwiftShaderTest, AttributeLocation)
+{
+	Initialize(3, false);
+
+	const std::string vs =
+		"#version 300 es\n"
+		"layout(location = 0) in vec4 a0;\n"   // Explicitly bound in GLSL
+		"layout(location = 2) in vec4 a2;\n"   // Explicitly bound in GLSL
+		"in vec4 a5;\n"                        // Bound to location 5 by API
+		"in mat2 a3;\n"                        // Implicit location
+		"in vec4 a1;\n"                        // Implicit location
+		"in vec4 a6;\n"                        // Implicit location
+		"out vec4 color;\n"
+		"void main()\n"
+		"{\n"
+		"   vec4 a34 = vec4(a3[0], a3[1]);\n"
+		"	gl_Position = a0;\n"
+		"   color = (a2 == vec4(1.0, 2.0, 3.0, 4.0) &&\n"
+		"            a34 == vec4(5.0, 6.0, 7.0, 8.0) &&\n"
+		"            a5 == vec4(9.0, 10.0, 11.0, 12.0) &&\n"
+		"            a1 == vec4(13.0, 14.0, 15.0, 16.0) &&\n"
+		"            a6 == vec4(17.0, 18.0, 19.0, 20.0)) ?\n"
+		"           vec4(0.0, 1.0, 0.0, 1.0) :\n"
+		"           vec4(1.0, 0.0, 0.0, 1.0);"
+		"}\n";
+
+	const std::string fs =
+		"#version 300 es\n"
+		"precision mediump float;\n"
+		"in vec4 color;\n"
+		"out vec4 fragColor;\n"
+		"void main()\n"
+		"{\n"
+		"	fragColor = color;\n"
+		"}\n";
+
+	ProgramHandles ph;
+	ph.program = glCreateProgram();
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	ph.vertexShader = glCreateShader(GL_VERTEX_SHADER);
+	const char* vsSource[1] = { vs.c_str() };
+	glShaderSource(ph.vertexShader, 1, vsSource, nullptr);
+	glCompileShader(ph.vertexShader);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+	GLint vsCompileStatus = 0;
+	glGetShaderiv(ph.vertexShader, GL_COMPILE_STATUS, &vsCompileStatus);
+	EXPECT_EQ(vsCompileStatus, GL_TRUE);
+
+	ph.fragmentShader = glCreateShader(GL_FRAGMENT_SHADER);
+	const char* fsSource[1] = { fs.c_str() };
+	glShaderSource(ph.fragmentShader, 1, fsSource, nullptr);
+	glCompileShader(ph.fragmentShader);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+	GLint fsCompileStatus = 0;
+	glGetShaderiv(ph.fragmentShader, GL_COMPILE_STATUS, &fsCompileStatus);
+	EXPECT_EQ(fsCompileStatus, GL_TRUE);
+
+	// Not assigned a layout location in GLSL. Bind it explicitly with the API.
+	glBindAttribLocation(ph.program, 5, "a5");
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	// Should not override GLSL layout location qualifier
+	glBindAttribLocation(ph.program, 8, "a2");
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	glAttachShader(ph.program, ph.vertexShader);
+	glAttachShader(ph.program, ph.fragmentShader);
+	glLinkProgram(ph.program);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	// Changes after linking should have no effect
+	glBindAttribLocation(ph.program, 0, "a1");
+	glBindAttribLocation(ph.program, 6, "a2");
+	glBindAttribLocation(ph.program, 2, "a6");
+
+	GLint linkStatus = 0;
+	glGetProgramiv(ph.program, GL_LINK_STATUS, &linkStatus);
+	EXPECT_NE(linkStatus, 0);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	float vertices[18] = { -1.0f,  1.0f, 0.5f,
+	                       -1.0f, -1.0f, 0.5f,
+	                        1.0f, -1.0f, 0.5f,
+	                       -1.0f,  1.0f, 0.5f,
+	                        1.0f, -1.0f, 0.5f,
+	                        1.0f,  1.0f, 0.5f };
+
+	float attributes[5][4] = { 1.0f, 2.0f, 3.0f, 4.0f,
+	                           5.0f, 6.0f, 7.0f, 8.0f,
+	                           9.0f, 10.0f, 11.0f, 12.0f,
+	                           13.0f, 14.0f, 15.0f, 16.0f,
+	                           17.0f, 18.0f, 19.0f, 20.0f };
+
+	GLint a0 = glGetAttribLocation(ph.program, "a0");
+	EXPECT_EQ(a0, 0);
+	glVertexAttribPointer(a0, 3, GL_FLOAT, GL_FALSE, 0, vertices);
+	glEnableVertexAttribArray(a0);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	GLint a2 = glGetAttribLocation(ph.program, "a2");
+	EXPECT_EQ(a2, 2);
+	glVertexAttribPointer(a2, 4, GL_FLOAT, GL_FALSE, 0, attributes[0]);
+	glVertexAttribDivisor(a2, 1);
+	glEnableVertexAttribArray(a2);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	GLint a3 = glGetAttribLocation(ph.program, "a3");
+	EXPECT_EQ(a3, 3);   // Note: implementation specific
+	glVertexAttribPointer(a3 + 0, 2, GL_FLOAT, GL_FALSE, 0, &attributes[1][0]);
+	glVertexAttribPointer(a3 + 1, 2, GL_FLOAT, GL_FALSE, 0, &attributes[1][2]);
+	glVertexAttribDivisor(a3 + 0, 1);
+	glVertexAttribDivisor(a3 + 1, 1);
+	glEnableVertexAttribArray(a3 + 0);
+	glEnableVertexAttribArray(a3 + 1);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	GLint a5 = glGetAttribLocation(ph.program, "a5");
+	EXPECT_EQ(a5, 5);
+	glVertexAttribPointer(a5, 4, GL_FLOAT, GL_FALSE, 0, attributes[2]);
+	glVertexAttribDivisor(a5, 1);
+	glEnableVertexAttribArray(a5);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	GLint a1 = glGetAttribLocation(ph.program, "a1");
+	EXPECT_EQ(a1, 1);   // Note: implementation specific
+	glVertexAttribPointer(a1, 4, GL_FLOAT, GL_FALSE, 0, attributes[3]);
+	glVertexAttribDivisor(a1, 1);
+	glEnableVertexAttribArray(a1);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	GLint a6 = glGetAttribLocation(ph.program, "a6");
+	EXPECT_EQ(a6, 6);   // Note: implementation specific
+	glVertexAttribPointer(a6, 4, GL_FLOAT, GL_FALSE, 0, attributes[4]);
+	glVertexAttribDivisor(a6, 1);
+	glEnableVertexAttribArray(a6);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	glUseProgram(ph.program);
+	glDrawArrays(GL_TRIANGLES, 0, 6);
+	EXPECT_GLENUM_EQ(GL_NONE, glGetError());
+
+	deleteProgram(ph);
+
+	unsigned char green[4] = { 0, 255, 0, 255 };
 	expectFramebufferColor(green);
 
 	EXPECT_GLENUM_EQ(GL_NONE, glGetError());